Erich Tilgner
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taxon links [up-->]Euphasmida [down<--]Neoptera Interpreting the tree
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Tilgner (2002)

Containing group: Neoptera


Phasmida are terrestrial, nocturnal, phytophagous insects found in nearly all temperate and tropical ecosystems (Günther, 1953). Scientists have described over 3,000 species (Bragg, 1995), yet this figure is uncertain since some taxon names are synonyms, and many new species have not been formally described.


Phasmida are variable in appearance, ranging from relatively generalized forms, to some that are wonderful mimics of sticks and/or leaves. They display varying degrees of brachyptery, and can be winged or wingless. The tarsi have three articles in Timema Scudder and five in other Phasmida. Cerci are composed of one article, except for adult males of Timema which have a lobe on the right cercus.

Sexual dimorphism is usually extreme: the males are smaller and more gracile than the females. Reproduction is typically sexual, but parthenogenesis occurs frequently. The egg capsule is distinctively shaped, possessing a lid called the operculum and a micropylar plate (Sellick, 1997). Eggs are large and oftentimes highly sculptured resembling plant seeds. They are laid singly, and are dropped, flicked, buried, glued to a surface, or riveted to a leaf. Some species that drop the eggs rely on ants to disperse them in a process analogous to myrmecochory (Windsor et al., 1996). The entire life cycle from egg to adult can take from several months to several years depending on the species.

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Egg capsule of Extatosoma popa. The nymph escapes by popping off the operculum. Photograph copyright © 2001, Erich Tilgner.

Phasmida possess several unique anatomical features that distinguish them from other Neoptera and indicate they are a monophyletic group (see Tilgner et al., 1999; 2002). For example:


Discussion of Phylogenetic Relationships

Timema is hypothesized to be the sister group to the remainder of Phasmida (Kristensen, 1975, Bradler 1999, Tilgner et al. 1999, Tilgner 2002), termed Euphasmida by Bradler (1999). Timema lack the autapomorphies of Euphasmida and possess traits found in other more distantly related Neoptera (Tilgner et al. 1999). 

Other Names for Phasmida


Bradler, S. 1999. The vomer of Timema Scudder, 1895 (Insecta: Phasmatodea) and its significance for phasmatodean phylogeny. Cour. Forschungstinst. Senckenb. 215:43-47.

Bragg, P.E. 1995. The Phasmid Database version 1.5. Phasmid Studies 3(2):41-42.

Günther, K. 1953. Über die taxonomische Gliederung und die geographische Verbreitung der Insektenordnung der Phasmatodea. Beiträge zur Entomol. 3:541-563.

Kristensen, N.P. 1975. Phylogeny of hexapod "orders." A critical review of recent accounts. Z. Zool. Syst. Evol.13:1-44.

Sellick, J.T.C. 1997. Descriptive terminology of the phasmid egg capsule, with an extended key to the phasmid genera based on egg structure. Syst. Entomol. 22:97-122.

Tilgner, E. H., Kiselyova, T. G., and McHugh, J. V. 1999. A morphological study of Timema cristinae Vickery with implications for the phylogenetics of Phasmida. Dtsche. Entomol. Z. 46(2):149-162.

Tilgner, E. H. 2002. Systematics of Phasmida. Ph.D. Dissertation, University of Georgia.

Wheeler, W.C. 1998. Sampling, groundplans, total evidence and the systematics of arthropods, In Fortey, R.A. & Thomas, R. H. (Eds.) Arthropod Relationships, Systematics Association special volume series 55, Chapman and Hall, London.

Windsor, D.M., Trapnell, D.W. and Amat, G. 1996. The egg capitulum of a neotropical walkingstick Calynda bicuspis, induces aboveground egg dispersal by the ponerine ant Ectatomma ruidum. J. Insect Behav. 9:353-362.

Information on the Internet

Title Illustrations
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Scientific Name Timema dorotheae
Location Hualapai Mountains, Arizona
Specimen Condition Live Specimen
Sex female
Image Use creative commons This media file is licensed under the Creative Commons Attribution License - Version 3.0.
Copyright © 2001 David R. Maddison
Scientific Name Heteropteryx dilatata
Specimen Condition Live Specimen
Sex Female
Life Cycle Stage nymph
Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0.
Copyright © 2001
About This Page

The author thanks Katja-Sabine Schulz and David R. Maddison for converting this page into the correct format for the Tree of Life. Tatiana Kiselyova kindly provided a review of this work.

University of Georgia, Athens, GA, USA

Correspondence regarding this page should be directed to Erich Tilgner at

Page: Tree of Life Phasmida. Phasmids. Authored by Erich Tilgner. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Tilgner, Erich. 2006. Phasmida. Phasmids. Version 26 November 2006. in The Tree of Life Web Project,

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